Palolo Valley (annual cultures) Palolo Valley (natural forest) Toro (monospecific planted shade trees/anoa dung) Temengor Forest Reserve (logger camp) Ba Be National Park (dung beetles) Danum Valley Field Centre (Riverine A) Sungkai Reserve Forest (secondary forest) The Ecological Register
Onthophagus rudis (scarab beetle): 7 samples
Standardised estimates are produced by subsampling and then extrapolating.
Raw richness may be higher when an abundance distribution is uneven.