Ursus (carnivore)

Abundance: 0.01 to 99.10% (median 1.52%)

Latitudinal range: 10.7° to 72.8°

Habitats: (172), boreal forest/taiga (6), desert/xeric shrubland (2), montane grassland (4), temperate broadleaf/mixed forest (17), temperate coniferous forest (5), temperate savanna (3), tropical/subtropical coniferous forest (2), tropical/subtropical dry broadleaf forest (8), tropical/subtropical moist broadleaf forest (1), tundra (1)

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view a larger map

Found in 221 samples

Afghanistan: Nuristan Province

Algeria: Ain Boucherit (Upper)

Armenia: Hovk-1 Cave (Unit 4)

Bhutan: Lamai Goempa Research Preserve

Bulgaria: Redaka II

Cambodia: Chhep Wildlife Sanctuary, Boh Dambang

Canada: Lindsay, Canadian National Railway (spring), Canadian National Railway (summer), Canadian National Railway (fall), Charlie Lake Cave (Zone IV), Learmonth Site, Peale Point House 2 (1), Keffer (Midden 57, screened), Keffer (Midden 57, floated), DjPp-8 (level B), Grassy Bay, Irving-Johnston, Caverne de la Mine (upper infill), Winefred Lake, EMEND site (retention level 100%, replicate 1), EMEND site (retention level 100%, replicate 2), EMEND site (retention level 100%, replicate 3), Algar

China: Wanglang National Nature Reserve, Shennongjia National Nature Reserve, Suojia Township, Nangunhe Nature Reserve, Guanyinshan Nature Reserve, Yanchiwan National Nature Reserve, Ma'anshan Cave (upper layer), Ma'anshan Cave (lower layer), Songhua River, Tengchong Section, Foping National Nature Reserve (2020), Kanas National Nature Reserve, Fanjingshan National Nature Reserve, Jinsitai Cave (Layer 7), Taosi, Zhoujiazhuang, Mocun Cave, Gouwan, Tianluoshan, Cunkong Cave, Lower Pubu Cave, Wuyun Cave, Mohui Cave, Ganxian Cave, Bailongdong, Yanlidong (Late Stage), Yanlidong (Early Stage)

Croatia: Northern Velebit National Park, Mujina Pecina (Level B), Mujina Pecina (Level D2)

Denmark: Mullerup (waste)

France: Grotte du Bison (Level I), Mas d'Azil, Les Auzières 2, Les Canalettes (Layer 2), Les Canalettes (Layer 3), Les Canalettes (Layer 4), Noisetier Cave, Coudoulous I (Faunal Unit II), Coudoulous I (Faunal Unit III), Coudoulous I (Faunal Units IV and V), Coudoulous I (Faunal Unit VI), Caune de l'Arago (Unit D), Saint-Vallier (LD3), Vergisson II, Romain-la-Roche, Lazaret (C II supérieur UA 13-17), Payre (ensemble G), Payre (ensemble F), Payre (ensemble D), Orgnac 3 (level 1), Tourville-la-Rivière (level D1), Soucy 1

Georgia: Ortvale Klde (late Middle Paleolithic), Ortvale Klde (early Upper Paleolithic), Dzudzuana Cave (lower area, Unit B), Kvabebi

Germany: Untermassfeld

India: Dachigam National Park

Iran: Lar Protected Area, Wezmeh Cave, Golestan National Park

Israel: Marj Rabba, Ubeidiya (Layer II-23)

Italy: Western Trento Province (2012), Caverna degli Orsi (sondage A, Unit 10), Caverna degli Orsi (sondage A, Unit 11.1), Caverna degli Orsi (sondage A, Unit 11.2), Riparo Biarzo (SU 5), Riparo Biarzo (SU 4 + 3B), Riparo Biarzo (SU 3A), Mondeval de Sora (SU 8), Mondeval de Sora (SU 31), Canale delle Acque Alte (Layer C2), La Sassa Cave, Pirro Nord 13, Poggetti Vecchi, Collecurti bonebed, Isernia La Pineta (3 colluvio level), Frantoio (Arda River), Grotta Cucgiliana, Poggio Rosso

Japan: Torihama

Laos: Tam Ngu Hao 2

Lebanon: Moghr el-Ahwal Cave 3 (Phases F & E)

Mexico: Ajos, Madera West, Schroeder Site (Structure 1), El Zurdo

Montenegro: Trlica (Bed 10), Trlica (Bed 5)

Morocco: Grotte des Rhinocéros (lower set), Grotte des Rhinocéros (upper set), Thomas Quarry Hominid Cave

Myanmar: Hukaung Valley Wildlife Sanctuary (Outside the Core), Kweenkoh, Mae Nyaw Kee South

North Korea: Chongphadae Cave

Poland: Borsuka Cave (Layer VI), Raj Cave (layer 4), Raj Cave (layer 6), Borsuka Cave (Layer III), Zabia Cave (Layer 17)

Portugal: Gruta de Figueira Brava (Area C)

Romania: Southern Transylvania, Noviodunum (TPF sector, Middle Ages)

Russia: Geographical Society Cave, Zapovednaya, Chagyrskaya Cave (Layer 5), Chagyrskaya Cave (Layer 6), Mezmaiskaya Cave (Layer 2A), Mezmaiskaya Cave (Layer 2B), Mezmaiskaya Cave (Layer 3), Cheremukhovo-1 (Bed 4), Imanay Cave (Pleistocene Layer 1), Imanay Cave (Pleistocene Layer 2), Imanay Cave (Pleistocene Layer 3), Imanay Cave (Pleistocene Layer 4), Khayrgas Cave - Layer 6 (1998–1999 pit), Khayrgas Cave - Layer 7 (1998–1999 pit), Byzovaya, Mezmaiskaya Cave (Layer 2)

Spain: Huéscar 1, Abric Romaní (Level Ja), Imanolen Arrobia, Cueva del Angel, Fuente Nueva 3 (layer 5), Bolomor Cave (Level IV), Venta Micena 4-I, Venta Micena 4-II, Barranc de la Boella (La Mina 2), Barranc de la Boella (El Forn 4), Barranco Léon, Gran Dolina (TD6.3), La Puebla de Valverde

Thailand: Khao Yai National Park, Khao Ang Rue Nai Wildlife Sanctuary, Cave of the Monk (Area 1), Ban Rai rockshelter, Tham Lod rockshelter (Unit 1), Tham Lod rockshelter (Unit 2), Tham Lod rockshelter (Unit 3)

Tunisia: Meninx

Turkey: Yaylacik Research Forest, Marmaris, Barhal Stream, Direkli Cave

Ukraine: Prolom II, Medzhibozh 1 (horizon 16a)

United Kingdom: Lynford Quarry, Balderton Sand and Gravel

United States: Salt Pond Mountain, Boulder County, North Fork of the Cache la Poudre River, North of the Michigamme Reservoir, Dalton Highway, Zesch Cave, Welsh Cave, Vess Cave, Papago Springs Cave, Haystack Cave (lower levels), Harrodsburg Crevice, Ripplemead, Bat Cave, Angel Site (Subdivision W10D), Wewukiyepuh, Fort Center Mound A, McLelland, Croley-Evans, Harris Creek, Garden Creek Mound No. 2, Hobson Site, Hale Farm, Dead Indian Creek, Prison Farm, Pine Hill, Otter Creek No. 2 (Zone 3), Cole Gravel Pit, Sachsen Cave Shelter, Robinson Cave, Polecat Creek (alluvial gravel), Schulze Cave (layer B), Schulze Cave (layer C1), Schulze Cave (layer C2), Natural Chimneys, Flagg Swamp Rockshelter (Archaic), Molybdenum Ridge (triggered detections), Molybdenum Ridge (time-lapse detections), Black Rapids Training Area (triggered detections), Bull Neck Swamp Research Forest, Smithsonian Conservation Biology Institute (logs, feature), West of US 287, Cumberland Cave

Vietnam: Duoi U'Oi, Lang Trang (Cave II, breccia 5), Coc Muoi

See also Ursus americanus, Ursus arctos, Ursus bibersoni, Ursus deningeri, Ursus dolinensis, Ursus etruscus, Ursus kudarensis, Ursus maritimus, Ursus minimus, Ursus sp., Ursus spelaeus, Ursus thibetanus, Ursus vitabilis