Ton Nga Chang Wildlife Sanctuary (primary forest) Ton Nga Chang Wildlife Sanctuary (secondary forest) Temengor Forest Reserve (logger camp) Ba Be National Park (dung beetles) Danum Valley Field Centre (Riverine A) Bangi Reserve Forest (secondary forest) Sungkai Reserve Forest (secondary forest) The Ecological Register
Paragymnopleurus (scarab beetle): 7 samples
Standardised estimates are produced by subsampling and then extrapolating.
Raw richness may be higher when an abundance distribution is uneven.