Jefferson City Las Gamas (EB) Las Gamas (PS) Blackbeard Island Cumberland Island El Hato del Volcan (large traps) El Hato del Volcan (small traps) San Jose de la Victoria Boca del Chajul East of Arapaho Prairie Arapaho Prairie Kibale National Park (K30) Minimum Critical Size of Ecosystems Project (contiguous forest) Welder Wildlife Refuge (open chaparral) Welder Wildlife Refuge (1985) Los Tuxtlas (coati dung/nocturnal) Viols 1 Viols 2 Kirrama State Forest (rain forest/intercept traps) Kirrama State Forest (open forest/intercept traps) Estación Experimental del Secretariado Uruguayo de la Lana (bovine dung) Estación Experimental del Secretariado Uruguayo de la Lana (ovine dung) Los Tuxtlas (continuous forest) Sierra de Baza (low) Sierra de Baza (medium) Sierra de Baza (high) Napu Valley (natural forest) Ton Nga Chang Wildlife Sanctuary (primary forest) Ton Nga Chang Wildlife Sanctuary (secondary forest) Fazenda Encanto (human feces baited) Fazenda Encanto (bovine meat baited) Paracou (roadside forest) Los Tuxtlas (howlers present) Los Tuxtlas (howlers absent) South of Busselton (dung bait) South of Busselton (carrion bait) Dhaka (cattle dung) Biological Dynamics of Forest Fragments Project (diurnal traps) Biological Dynamics of Forest Fragments Project (nocturnal traps) Jalcomulco Real Minero Thirunelly Forests Chiruisla Station Viçosa (cow dung) Viçosa (horse dung) Ilha Solteira Palolo Valley (annual cultures) Maputo Special Reserve (reserve grassland) Kurukshetra Charles Darwin Ecological Refuge (open habitat) Charles Darwin Ecological Refuge (forest habitat) Cikrikci (2005) Lola Field Station (savanna) Lola Field Station (forest) North of Caldwell (open pasture, 1979) Centro Cientifico de Rio Palenque (May 1975) Centro Cientifico de Rio Palenque (February 1976) Centro Cientifico de Rio Palenque (July 1976) Hluhluwe Game Reserve Santa Rosa Salamanca (spring) Campos Novos (feces baited) Campos Novos (carrion baited) Elí Farm (howler monkey dung baited) Elí Farm (woolly monkey dung baited) Nouragues Biological Field Station (hour 21) Barreiro Protection Area (cerrado) Barreiro Protection Area (forest) Verdon Valley M[1] (July) Mejerda Valley San Martín Volcano (forest fragments) Piritáma II Ferret Valley (shrubbery) Fazenda Canchim Manga Rosa Morro do Elefante Morro do Cerrito Campus da Universidade Federal de Santa Maria Bagé (Eragrostis plana) Bagé (Eucalyptus spp.) Ankasa Conservation Area (unlogged forest) Wongabel State Forest Hutcheson Memorial Forest disturbed field (June/July) Hutcheson Memorial Forest old growth forest (June/July) Rutgers University Bovine Farm (June/July) Purple Springs Grazing Reserve A Purple Springs Grazing Reserve B Purple Springs Grazing Reserve C Chiusa Pesio Lake City Research Center Site A Lake City Research Center Site B Northeast of Bundaberg (1990 - 1991) Vicente Guerrero (IF) Reserva Natural La Planada (primary forest site 1) Nouragues Research Station (2003 FIT/creek) Animal Disease Research Institute (1995) Sierra de Salinas and Sierra de Onil (cattle dung, May) Sierra de Salinas and Sierra de Onil (rabbit dung, May) Upper Hunter Catchment (mature native woodland) Odivelas County Cork and County Tipperary (organic farms) Ohdaigahara Site A Ohdaigahara Site B Ohdaigahara Site C Brookings County (cattle manure) Brookings County (sheep manure) Kutari Toro (natural forest/anoa dung) Ipassa Reserve (large rainy season) Majawanga (bush site 1) Parque Nacional Noel Kempff Mercado (1997 forest samples) Singhori Wildlife Sanctuary (mixed forests site 2) Atariya Chhaparwa Lambir Hills National Park (isolated primary forests) Donggu Tongling Decha Laboratorio del Desierto (dung pats) Laboratorio del Desierto (pitfall trap site 1) El Jaralillo Cattle Ranch (July) Serranía de Coroza (dry forest, rainy season) Refugio del Oso de Anteojos Northeastern Tochigi Prefecture Kibale National Park (Ngogo) Kibale National Park (K14) Kibale National Park (K15) Kibale National Park (Pinus plantation) Kibale National Park (Cupressus plantation) Napu Valley (young secondary forest) Napu Valley (agroforestry ecosystem) Napu Valley (annual culture) Palolo Valley (natural forest) Maputo Special Reserve (reserve farm) San Martín Volcano (forest-pasture edges) San Martín Volcano (pastures) Ankasa Conservation Area (plantation) Vicente Guerrero (PE) Reserva Natural La Planada (secondary forest site 1) Reserva Natural La Planada (pasture site 1) Reserva Natural La Planada (cropland site 1) Upper Hunter Catchment (unplanted) Toro (monospecific planted shade trees/anoa dung) Toro (open areas/anoa dung) Majawanga (field site 4) Singhori Wildlife Sanctuary (agricultural lands site 4) Lambir Hills National Park (rubber garden) Serranía de Coroza (livestock use area, rainy season) Minimum Critical Size of Ecosystems Project (clearcuts) Minimum Critical Size of Ecosystems Project (1 ha forest fragment) Minimum Critical Size of Ecosystems Project (10 ha forest fragment) Los Tuxtlas (mosaic habitat) Los Tuxtlas (forest fragments) Paracou (interior forest) San Fernando (tropical deciduous forest) San Fernando (pasture with Acacia shrubs) Tepexco (June) Temengor Forest Reserve (logger camp) La Belgique (old secondary forest) La Belgique (sleeping sites) Highland Road Observatory Park (March) Missouri Ozark Forest Ecosystem Project (no harvest management) Floresta Nacional Contendas do Sincorá (dry season) Floresta Nacional Contendas do Sincorá (wet season) Vilela Farm Area de Proteção Ambiental de Cafuringa (feces-baited) Guajará Mirim e Nova Mamoré (baço bovino) Nouragues Research Station (November 2007) Amazone Nature Lodge (February 2009) Mont Grand Matoury National Nature Reserve (February 2009) Ba Be National Park (dung beetles) Mo Singto Long Term Ecological Research plot (macaque) Mo Singto Long Term Ecological Research plot (barking deer) Mo Singto Long Term Ecological Research plot (sambar deer) Mo Singto Long Term Ecological Research plot (elephant) Mo Singto Long Term Ecological Research plot (control) Mata Seca State Park (December) Skukuza (control) Baiyinxile Ranch (horse manure) Davies Creek (dung beetles, early wet season) Davies Creek (dung beetles, late wet season) Davies Creek (dung beetles, early dry season) Davies Creek (dung beetles, late dry season) Danum Valley Field Centre (Riverine A) Torre Paterno Makerere University Biological Field Station (dung beetles) Dagmarmara (600 m, 2004) Sialkot Jhelum (pastures) Jhelum (croplands) Bangi Reserve Forest (secondary forest) Sungkai Reserve Forest (secondary forest) Shiraoi (horse dung) Shiraoi (cattle dung) Mukawa (horse dung) Mukawa (cattle dung) The Ecological Register: Scarab Beetles
scarab beetles: 196 samples
Standardised estimates are produced by subsampling and then extrapolating.
Raw richness may be higher when an abundance distribution is uneven.