Abundance: 0.01 to 71.43% (median 0.60%)

Latitudinal range: 39.7° to 63.8°

Habitats: (20), boreal forest/taiga (4), montane grassland (3)

Found in 27 samples

Canada: Charlie Lake Cave (Zone IV), EMEND site (retention level 100%, replicate 1), EMEND site (retention level 100%, replicate 2), Algar

China: Kanas National Nature Reserve, Jinsitai Cave (Layer 7)

France: Vergisson II, Romain-la-Roche

Mongolia: Otson Tsokhio and Zuun Shovkh

Poland: Borsuka Cave (Layer VI), Zabia Cave (Layer 19), Zabia Cave (Layer 18), Zabia Cave (Layer 17), Zabia Cave (Layer 15), Zabia Cave (Layer 13), Zabia Cave (Layer 12), Zabia Cave (Layer 9)

Russia: Geographical Society Cave, Bol'shoj Naryn (Paleosol Layers), Imanay Cave (Pleistocene Layer 1), Imanay Cave (Pleistocene Layer 2), Imanay Cave (Pleistocene Layer 3), Imanay Cave (Pleistocene Layer 4)

United States: Molybdenum Ridge (triggered detections), Molybdenum Ridge (time-lapse detections), Black Rapids Training Area (triggered detections), Cumberland Cave

See also Gulo gulo, Gulo schlosseri, Gulo sp.