Abundance: 0.02 to 7.33% (median 0.55%)

Latitudinal range: 32.2° to 51.4°

Habitats: (21), Mediterranean woodland (1), temperate broadleaf/mixed forest (3)

Found in 25 samples

Czech Republic: Merkur−North (Early Miocene), Mokrá-Western Quarry (Early Miocene)

France: Coudoulous I (Faunal Unit II) (Middle Pleistocene), Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Romain-la-Roche (Middle Pleistocene), Lazaret (C II supérieur UA 13-17) (Middle Pleistocene)

Germany: Hambach 6C (Middle Miocene), Griesbeckerzell 1a (Middle Miocene)

Greece: Lake Prespa National Park

Iran: Wezmeh Cave (Late Pleistocene)

Italy: Poggi di Prata, Monte Argentario (Early Pleistocene), Monticino Quarry (Late Miocene)

Jordan: Dibbeen Nature Reserve

Kazakhstan: Baikadam (Middle Miocene)

Montenegro: Great Ulcinj Beach

Poland: Zabia Cave (Layer 21) (Early Pleistocene), Zabia Cave (Layer 17) (Early Pleistocene), Zabia Cave (Layer 15) (Early Pleistocene)

Romania: Subpiatra 2/1R (Middle Miocene)

Spain: Caló den Rafelino (Early Pliocene), Puerto de la Cadena (Early Pliocene)

Ukraine: Grot Skeliastyi (III-3-x) (Late Pleistocene)

See also Vipera ammodytes, Vipera aspis, Vipera berus, Vipera lebetina, Vipera natiensis, Vipera palaestinae, Vipera s.l., Vipera sp., Vipera sp. 2, Vipera sp. European vipers group, Vipera sp. Oriental vipers group, Vipera sp. Vipera aspis complex, Vipera ursinii