Abundance: 0.07 to 28.57% (median 1.43%)

Latitudinal range: 0.1° to 51.9°

Habitats: (15) only

Found in 15 samples

Ecuador: La Chimba (Holocene)

Egypt: Gebel Silsila 2B-II (Silsilian) (Late Pleistocene)

Ethiopia: Andalee (Middle Pleistocene)

Germany: Echzell (Early Miocene)

Greece: Ano Metochi-2 (Late Miocene), Ano Metochi-3 (Late Miocene), Pikermi (Late Miocene)

Lebanon: Moghr el-Ahwal Cave 3 (Phases F & E) (Late Pleistocene)

Niger: Garumele (context 2) (Holocene)

Russia: Kuznetsovka (Middle Pleistocene), Muhkai II (layer 80) (Early Pleistocene)

Saudi Arabia: Ti’s al Ghadah (Middle Pleistocene)

United States: Crystal Ball Cave (Late Pleistocene), Ripplemead (Late Pleistocene), McIntosh (Holocene)

See also Squamata indet.