Habitats: temperate broadleaf/mixed forest (8)
onlyFound in 15 samples
China: Kajun Village (MK-1) (Early Oligocene), Kajun Village (MK-3) (Late Eocene)
Denmark: Bolderslev Skov
Greece: Dadia-Lefkimi-Soufi (nest-sites), Akrocheiras (Early Miocene)
Hungary: Wind Brickyard (Late Oligocene)
India: Binsar Wildlife Sanctuary
Japan: Chichibu Mountains, University of Tokyo Hokkaido Forest, Ken-Marubi Lava Flow
United States: Central West Virginia, Yosemite National Park, USNM locality no. 41352 (Early Eocene), East Brook (Middle Eocene), West Brook (Middle Eocene)
See also Alnus ferdinandicoburgii, Alnus firma, Alnus glutinosa, Alnus hirsuta, Alnus incana, Alnus nepalensis, Alnus oligocaenica, Alnus rhombifolia, Alnus rugosa, Alnus sp., Alnus sp. 1, Alnus sp. HER22, Alnus sp. HER38